| |||
|---|---|---|---|
This list of fossil fishes described in 2019 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes of every kind that were described during the year 2019, as well as other significant discoveries and events related to paleoichthyology that occurred in 2019.
New taxa
Jawless vertebrates
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Archegonaspis bashkirica[1] |
Sp. nov |
Valid |
Märss |
A member of the family Cyathaspididae. |
||||
|
Sp. nov |
Valid |
Märss |
Tabuska Beds |
A member of the family Cyathaspididae. |
||||
|
Jiaoyu[2] |
Gen. et sp. nov |
Valid |
Liu et al. |
Kezirtag formation |
A member of Galeaspida of uncertain phylogenetic placement. The type species is J. imperfectus. |
|||
|
Parathelodus liaokuoensis[3] |
Sp. nov |
Valid |
Cui et al. |
A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020. |
||||
|
Parathelodus wangi[3] |
Sp. nov |
Valid |
Cui et al. |
A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020. |
||||
|
Parathelodus xitunensis[3] |
Sp. nov |
Valid |
Cui et al. |
A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020. |
||||
|
Gen. et sp. nov |
Valid |
Miyashita et al. |
A hagfish. The type species is T. tapirostrum. |
|||||
|
Tolypelepis bedovensis[1] |
Sp. nov |
Valid |
Märss |
Late Silurian |
Krasnaya Bukhta Formation |
A member of the family Cyathaspididae. |
||
|
Xiyuaspis[2] |
Gen. et comb. nov |
Junior homonym |
Liu et al. |
Tataertag Formation |
A member of Galeaspida belonging to the group Hanyangaspidida and to the family Xiushuiaspidae. The type species is "Nanjiangaspis" zhangi Lu et al. (2007). The generic name is preoccupied by Xiyuaspis Zhang (1985); Shan et al. (2023) coined a replacement name Xiyuichthys.[5] |
|||
Placoderms
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Sp. nov |
Valid |
Downs et al. |
An antiarch in Asterolepis |
|||||
|
Colombialepis[7] |
Gen. et sp. nov |
Valid |
Olive et al. |
Late Devonian |
A member of Antiarchi, superfamily Asterolepidoidei. Genus includes new species C. villarroeli. |
|||
|
Colombiaspis[7] |
Gen. et sp. nov |
Valid |
Olive et al. |
Late Devonian |
A member of Arthrodira, family Groenlandaspidae. Genus includes new species C. rinconensis. |
|||
|
Meeksiella[8] |
Gen. et sp. nov |
Valid |
Trinajstic et al. |
Snetnya Gora Beds |
A member of Ptyctodontida. The type species is "Chelyophorus" pskovensis Obruchev (1947). |
|||
.svg.png.webp){kind=small}
Acanthodians
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Sp. nov |
Valid |
Den Blaauwen, Newman & Burrow |
Mey Flagstone Formation |
|||||
|
Serradentus[10] |
Gen. et sp. nov |
Newman, Burrow & Den Blaauwen |
Tordalen Formation |
A member of Ischnacanthiformes. The type species is S. armstrongi. |
||||
Cartilaginous fishes
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et sp. nov |
Valid |
Stumpf & Kriwet |
A member of Rajiformes of uncertain phylogenetic placement. The type species is A. grimmenensis. |
|||||
|
Asflapristis[12] |
Gen. et sp. nov |
Valid |
Villalobos-Segura et al. |
Akrabou Formation |
A member of Sclerorhynchoidei belonging to the family Ptychotrygonidae. Genus includes new species A. cristadentis. |
|||
|
Sp. nov |
Valid |
Case et al. |
Bladen Formation |
A member of the family Ginglymostomatidae. |
||||
|
Carcharhinus mancinae[14] |
Sp. nov |
Valid |
Ebersole, Cicimurri & Stringer |
A species of Carcharhinus. |
||||
|
Carcharhinus underwoodi[15] |
Sp. nov |
Valid |
Samonds et al. |
Mahajanga Basin |
A species of Carcharhinus. |
|||
|
Carcharias dominguei[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
A species of Carcharias. |
||||
|
Centrosqualus mustardi[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A member of the family Squalidae. |
||
|
Chlamydoselachus balli[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A species of Chlamydoselachus. |
||
|
Sp. nov |
Valid |
Hoganson, Erickson & Holland |
||||||
|
Sp. nov |
Valid |
Shimada & Everhart |
A mackerel shark belonging to the family Pseudoscapanorhynchidae. |
|||||
|
Dasyatis northdakotaensis[17] |
Sp. nov |
Valid |
Hoganson, Erickson & Holland |
A species of Dasyatis. |
||||
|
Diprosopovenator[19] |
Gen. et sp. nov |
Valid |
Stumpf, Scheer & Kriwet |
Hesseltal Formation |
A member of Carcharhiniformes belonging to the new family Pseudoscyliorhinidae. The type species is D. hilperti. |
|||
|
Gen. et sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A member of the family Chlamydoselachidae. The type species is D. garethi. |
|||
|
Florenceodon[16] |
Gen. et sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A member of Carcharhiniformes belonging to the new family Florenceodontidae. Genus includes new species F. johnyi. |
||
|
Gen. et sp. nov |
Valid |
Gates, Gorscak & Makovicky |
A carpet shark. The type species is G. nordquistae. |
|||||
|
Glencartius[21] |
Gen. et comb. nov |
Valid |
Ginter & Skompski |
A member of Ctenacanthiformes of uncertain phylogenetic placement. The type species is "Ctenacanthus" costellatus Traquair (1884). |
||||
|
Gydoselache[22] |
Gen. et sp. nov |
Valid |
Maisey et al. |
Gydo Formation |
A cartilaginous fish of uncertain phylogenetic placement, related to Pucapampella and assigned to the new family Pucapampellidae. The type species is G. oosthuizeni. |
|||
|
Gen. et sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A member of the family Dalatiidae. Genus includes new species H. wardi. |
|||
|
Hypolophites beckeri[23] |
Sp. nov |
Valid |
Maisch |
A member of Myliobatiformes. |
||||
|
Isogomphodon aikenensis[24] |
Sp. nov |
Valid |
Cicimurri & Knight |
Dry Branch Formation |
A relative of the daggernose shark. |
|||
|
Kimmerobatis[25] |
Gen. et sp. nov |
Valid |
Underwood & Claeson |
A member of Batoidea related to Spathobatis. Genus includes new species K. etchesi. |
||||
|
Gen. et sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A member of Hexanchiformes belonging to the new family Komoksodontidae. Genus includes new species K. kwutchakuth. |
|||
|
Gen. et sp. nov |
Valid |
Marramà et al. |
Monte Bolca Konservat-Lagerstätte |
A member of Myliobatiformes belonging to the superfamily Dasyatoidea. The type species is L. aenigmatica. |
||||
|
Sp. nov |
Valid |
Otero |
||||||
|
"Myliobatis" foxhillsensis[17] |
Sp. nov |
Valid |
Hoganson, Erickson & Holland |
An eagle ray. |
||||
|
Odontorhytis bahariensis[28] |
Sp. nov |
Valid |
Salame & Asan |
A member of Neoselachii of uncertain phylogenetic placement. |
||||
|
Ottangodus[29] |
Gen. et sp. nov |
Valid |
Popov, Delsate & Felten |
Luxembourg-French border |
A chimaera belonging to the family Callorhinchidae. Genus includes new species O. lotharingiae. |
|||
|
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
||||
|
Pastinachus kebarensis[30] |
Sp. nov |
Valid |
Adnet et al. |
A cowtail stingray of Pastinachus. Announced in 2018; the final version of the article naming it was published in 2019. |
||||
|
Pristiophorus humboldti[31] |
Sp. nov |
Valid |
Villafaña et al. |
Early Miocene |
A species of Pristiophorus. |
|||
|
Pristiophorus pricei[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A species of Pristiophorus. |
||
|
Pristiophorus smithi[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A species of Pristiophorus. |
||
|
Protocentrophorus steviae[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A member of the family Centrophoridae. |
||
|
Protoheptranchias[16] |
Gen. et sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A cow shark. Genus includes new species P. lowei. |
||
|
Sp. nov |
Valid |
Carrillo-Briceño, Parra & Luque |
Early Cretaceous (late Barremian–early Aptian) |
A mackerel shark belonging to the family Pseudoscapanorhynchidae. |
||||
|
Pseudabdounia[14] |
Gen. et comb. nov |
Valid |
Ebersole, Cicimurri & Stringer |
A requiem shark. The type species is "Galeocerdo" recticonus Winkler (1874); genus also includes "Galeorhinus recticonus" claibornensis White (1956) (raised to the rank of a separate species Pseudabdounia claibornensis). |
||||
|
Sp. nov |
Valid |
Villalobos‐Segura, Underwood & Ward |
||||||
|
Rhinoscymnus clarki[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A member of the family Somniosidae. |
||
|
Gen. et sp. et comb. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous to Miocene |
Angostura Formation |
|
A member of the family Chlamydoselachidae. The type species is R. ludvigseni; genus also includes "Chlamydoselachus" bracheri Pfeil (1983), "Chlamydoselachus" fiedleri Pfeil (1983), "Chlamydoselachus" thomsoni Richter & Ward (1990), "Chlamydoselachus" goliath Antunes & Cappetta (2002), "Chlamydoselachus" keyesi Mannering & Hiller (2008), "Chlamydoselachus" tatere Consoli (2008) and "Chlamydoselachus" landinii Carrillo-Briceño, Aguilera & Rodriguez (2014). |
||
|
Scymnodalatias kazenobon[34] |
Sp. nov |
Valid |
Nishimatsu & Ujihara |
Middle Miocene |
Yatsuo Group |
A species of Scymnodalatias. |
||
|
Squaliodalatias savoiei[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A member of the family Dalatiidae. |
||
|
Squalus nicholsae[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A spurdog. |
||
|
Synechodus dereki[16] |
Sp. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A member of the family Palaeospinacidae. |
||
|
Gen. et comb. nov |
Valid |
Marramà et al. |
A whiptail stingray belonging to the subfamily Neotrygoninae; a new genus for "Raja" muricata Volta (1796). |
|||||
|
Tlalocbatos[36] |
Gen. et sp. nov |
Valid |
Brito, Villalobos-Segura & Alvarado-Ortega |
A member of Batoidea related to the banjo rays. Genus includes new species T. applegatei. |
||||
|
Gen. et comb. nov |
Valid |
Cappetta, Morrison & Adnet |
Late Cretaceous (Campanian) |
Northumberland Formation |
A cow shark. Genus includes "Notidanus" dentatus Woodward (1886). |
|||
.svg.png.webp){kind=small}
Ray-finned fishes
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Acronuroides[37] |
Gen. et sp. nov. |
Valid |
Bannikov, Carnevale & Tyler |
Monte Bolca locality |
A member of Percomorphacea of uncertain phylogenetic placement. The type species is A. eocaenicus. |
|||
|
Acropoma aurora[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Acropoma. |
||||
|
Ambassis simesi[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Ambassis. |
||||
|
Sp. nov |
Valid |
Schwarzhans |
A member of the family Ophidiidae. |
|||||
|
Antigonia artata[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Antigonia. |
||||
|
Antimyctophum[38] |
Gen. et comb. nov |
Valid |
Schwarzhans |
A lanternfish. The type species is "Scopelus" konganaruensis Frost (1933). |
||||
|
Archoplites langrellorum[39] |
Sp. nov |
Valid |
Van Tassell & Smith |
A relative of the Sacramento perch. |
||||
|
Gen. et sp. nov |
Valid |
Romano et al. |
A member of the family Ptycholepidae. Genus includes new species A. occidentalis. |
|||||
|
Argyripnus oamaruensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Argyripnus. |
||||
|
Argyripnus rapahoensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Argyripnus. |
||||
|
Arnoglossus lautus[38] |
Sp. nov |
Valid |
Schwarzhans |
A scaldfish. |
||||
|
Arnoglossus purus[38] |
Sp. nov |
Valid |
Schwarzhans |
A scaldfish. |
||||
|
Artediellichthys candelabrum[41] |
Sp. nov |
Valid |
Nazarkin |
Agnevo Formation |
A relative of the blackfin hookear sculpin. |
|||
|
Artediellus simplex[41] |
Sp. nov |
Valid |
Nazarkin |
Agnevo Formation |
A species of Artediellus. |
|||
|
Aseraggodes hudsoni[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Aseraggodes. |
||||
|
Auchenoceros simplex[38] |
Sp. nov |
Valid |
Schwarzhans |
A relative of the ahuru. |
||||
|
Aulichthys miocaenicus[42] |
Sp. nov |
Valid |
Nazarkin |
Agnevo Formation |
An extinct tubenose. |
|||
|
Ausonasynodus[43] |
Gen. et sp. nov |
Valid |
Carnevale et al. |
Late Eocene |
A lizardfish. Genus includes new species A. almerai. |
|||
|
Awamoa[38] |
Gen. et comb. et sp. nov |
Valid |
Schwarzhans |
A southern sandfish. The type species is "Leptoscopus" progressus Schwarzhans (1980); genus also includes "Leptoscopus" iocosus Schwarzhans (1980) and a new species A. kaawa. |
||||
|
Balistes vegai[44] |
Sp. nov |
Valid |
Viñola Lopez, Carr & Lorenzo |
A species of Balistes. |
||||
|
Barathronus nielseni[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Barathronus. |
||||
|
Barbalepis[45] |
Gen. et comb. nov |
Valid |
Olive, Taverne & López-Arbarello |
A member the family Coccolepididae; a new genus for "Coccolepis" macroptera Traquair. |
||||
|
Bathycongrus waihaoensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Congridae. Originally described as a species of Bathycongrus, but subsequently transferred to the genus Smithconger.[46] |
||||
|
Bathygadus waiohaensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Bathygadus. |
||||
|
Bembrops aequiformis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Bembrops. |
||||
|
Bembrops grenfelli[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Bembrops. |
||||
|
Benthosema minutum[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Benthosema. |
||||
|
Bidenichthys struthersi[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Bidenichthys. |
||||
|
Blachea semeniformis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Blachea. |
||||
|
Bonapartia altidorsalis[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Gonostomatidae. |
||||
|
Brosmophyciops tongarewae[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Brosmophyciops. |
||||
|
Burguklia minichorum[47] |
Sp. nov |
Valid |
Bakaev & Kogan |
An early ray-finned fish. Announced in 2019; the final version of the article naming it was published in 2020. |
||||
|
Cabindachanos[48] |
Gen. et sp. nov |
Valid |
Taverne et al. |
A member of the family Chanidae. The type species is C. dartevellei. |
||||
|
Callionymus hakatarameaensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Callionymus. |
||||
|
Callionymus triquetrus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Callionymus. |
||||
|
Gen. et sp. nov |
Valid |
Romano et al. |
A stem-neopterygian belonging to the family Parasemionotidae. Genus includes new species C. argentus. |
|||||
|
Cavinichthys[49] |
Gen. et sp. nov |
Valid |
Taverne & Capasso |
Limestones of Pietraroja |
A member of Crossognathiformes belonging to the family Pachyrhizodontidae. The type species is C. pachylepis. |
|||
|
Cephalopholis? aotearoa[38] |
Sp. nov |
Valid |
Schwarzhans |
Possibly a species of Cephalopholis. |
||||
|
Cepola ambifaria[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Cepola. |
||||
|
Ceratoscopelus richardsoni[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Ceratoscopelus. |
||||
|
Champsodon timaruensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Champsodon. |
||||
|
Chaunax bucculentus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Chaunax. |
||||
|
Chaychanus[50] |
Gen. et sp. nov |
Valid |
Cantalice Severiano, Alvarado Ortega & Bellwood |
A member of the family Pomacentridae. Genus includes new species C. gonzalezorum. Announced in 2019; the final version of the article naming it was published in 2020. |
||||
|
Chrionema salebrosa[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Chrionema. |
||||
|
Coelorinchus divulgatus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Coelorinchus. |
||||
|
Coelorinchus pakaurangiensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Coelorinchus. |
||||
|
Coelorinchus preaustralis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Coelorinchus. |
||||
|
Sp. nov |
Valid |
Li et al. |
Falang Formation |
A member of Perleidiformes. |
||||
|
Conger davidsmithi[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Conger. |
||||
|
Conger tokoroa[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Conger. |
||||
|
Gen. et sp. nov |
Valid |
Taverne |
Stanleyville Formation |
A member of Pycnodontiformes belonging to the superfamily Coccodontoidea. The type species is C. cornutus. |
||||
|
Costapycnodus[53] |
Gen. et comb. nov |
Valid |
Taverne, Capasso & Del Re |
Early Cretaceous (late Hauterivian-early Barremian) |
A member of the family Pycnodontidae. The type species is "Coelodus" costae Heckel (1856). |
|||
|
Danaphos glomerosus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Danaphos. |
||||
|
Diaphus audax[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Diaphus. |
||||
|
Diaphus caurus[54] |
Sp. nov |
Valid |
Schwarzhans & Ohe |
Hijikata Formation |
A species of Diaphus. |
|||
|
Diaphus endoi[54] |
Sp. nov |
Valid |
Schwarzhans & Ohe |
Shinzato Formation |
A species of Diaphus. |
|||
|
Diaphus exilis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Diaphus. |
||||
|
Diaphus grebneffi[54] |
Sp. nov |
Valid |
Schwarzhans & Ohe |
Probably Piacenzian |
Nakosi Formation |
A species of Diaphus. |
||
|
Diaphus huatau[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Diaphus. |
||||
|
Diaphus kaiparaensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Diaphus. |
||||
|
Diaphus kakegawaensis[54] |
Sp. nov |
Valid |
Schwarzhans & Ohe |
A species of Diaphus. |
||||
|
Diaphus manneringi[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Diaphus. |
||||
|
Diaphus mirus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Diaphus. |
||||
|
Diaphus nafpaktitisi[54] |
Sp. nov |
Valid |
Schwarzhans & Ohe |
Nobori Formation |
A species of Diaphus. |
|||
|
Diaphus noboriensis[54] |
Sp. nov |
Valid |
Schwarzhans & Ohe |
Pliocene (Piacenzian) and Pleistocene (Gelasian) |
Dainichi Formation |
A species of Diaphus. |
||
|
Diaphus tenax[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Diaphus. |
||||
|
Dibranchus kakahoensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Dibranchus. |
||||
|
Diogenichthys rangiauriensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Diogenichthys. |
||||
|
Dipulus certus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Dipulus. |
||||
|
Dolichopteryx iustus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Dolichopteryx. |
||||
|
Echiodon teres[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Echiodon. |
||||
|
Electrona subasperoides[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Electrona. |
||||
|
Ellimma longipectoralis[55] |
Sp. nov |
Valid |
Polck et al. |
Barra Velha Formation |
A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Paraclupeidae. Announced in 2019; the final version of the article naming it was published in 2020. |
|||
|
Emmelichthys tennysoni[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Emmelichthys. |
||||
|
Encheliophis strigosus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Encheliophis. |
||||
|
Sp. nov |
Valid |
Schwarzhans |
A lanternfish. |
|||||
|
Sp. nov |
Valid |
Schwarzhans |
A lanternfish. |
|||||
|
Sp. nov |
Valid |
López-Arbarello et al. |
Meride Limestone |
A member of Semionotiformes. |
||||
|
Sp. nov |
Valid |
López-Arbarello et al. |
Meride Limestone |
A member of Semionotiformes. |
||||
|
Sp. nov |
Valid |
López-Arbarello et al. |
Meride Limestone |
A member of Semionotiformes. |
||||
|
Epaelops[57] |
Gen. et sp. nov |
Valid |
Alves, Alvarado-Ortega & Brito |
A member of Elopiformes. Genus includes new species E. martinezi. Announced in 2019; the final version of the article naming it was published in 2020. |
||||
|
Epigonus aquilonius[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Epigonus. |
||||
|
Epigonus opoitiensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Epigonus. |
||||
|
Etrumeus carnatus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Etrumeus. |
||||
|
Evynnis abax[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Sparidae. |
||||
|
Flagellipinna[58] |
Gen. et sp. nov |
Valid |
Cawley & Kriwet |
A member of Pycnodontiformes belonging to the family Pycnodontidae. The type species is F. rhomboides. |
||||
|
Forsterygion jawadi[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Forsterygion. |
||||
|
Gadiculus adversus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Gadiculus. |
||||
|
Gadiculus pereawa[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Gadiculus. |
||||
|
Galeichthys ohei[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Galeichthys. |
||||
|
Gilmourella[59] |
Gen. et sp. nov. |
Valid |
Carnevale & Bannikov |
Monte Bolca locality |
A member of Callionymoidei. The type species is G. minuta. |
|||
|
Gnathophis araroa[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Gnathophis. |
||||
|
Gonostoma retunsum[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Gonostoma. |
||||
|
Grahamichthys frigophila[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Thalasseleotrididae. |
||||
|
Guus[60] |
Gen. et comb. nov. |
Valid |
Bannikov |
Monte Bolca locality |
A member of the family Tortonesidae. The type species is "Gobius" microcephalus Agassiz (1839). |
|||
|
Sp. nov |
Valid |
Gouiric-Cavalli, Remírez & Kriwet |
Early Cretaceous (Valanginian–early Hauterivian) |
|||||
|
Harpagifer? morgansi[38] |
Sp. nov |
Valid |
Schwarzhans |
Possibly a species of Harpagifer. |
||||
|
Hemerocoetes pukunati[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Hemerocoetes. |
||||
|
Hemerocoetes whiroki[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Hemerocoetes. |
||||
|
Hiascoactinus[62] |
Gen. et sp. nov |
Valid |
Kim et al. |
Amisan Formation |
A member of Redfieldiiformes. Genus includes new species H. boryeongensis. Announced in 2019; the final version of the article naming it was published in 2020. |
|||
|
Hintonia robertsi[38] |
Sp. nov |
Valid |
Schwarzhans |
A lanternfish. |
||||
|
Howella monodens[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Howella. |
||||
|
Hygophum acutiventris[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Hygophum. |
||||
|
Ichthyscopus pukeuriensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Ichthyscopus. |
||||
|
Klincigobius[63] |
Gen. et comb. et sp. nov |
Valid |
Bradić-Milinović, Ahnelt & Schwarzhans in Bradić-Milinović et al. |
Early Miocene |
A member of the family Gobiidae. The type species is "Gobius" serbiensis Gaudant (1998); genus also includes new species K. andjelkovicae. |
|||
|
Kradimus[64] |
Gen. et sp. nov |
Valid |
Veysey, Brito & Martill |
Akrabou Formation |
A member of Crossognathiformes. Genus includes new species K. asflaensis. Announced in 2019; the final version of the article naming it was published in 2020. |
|||
|
Lactarius primigenius[38] |
Sp. nov |
Valid |
Schwarzhans |
A relative of the false trevally. |
||||
|
Lactarius pusillus[38] |
Sp. nov |
Valid |
Schwarzhans |
A relative of the false trevally. |
||||
|
Laeops undulatus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Laeops. |
||||
|
Lampanyctodes transitivus[38] |
Sp. nov |
Valid |
Schwarzhans |
A lanternfish. |
||||
|
Lampanyctus popoto[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Lampanyctus. |
||||
|
Lampanyctus profestus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Lampanyctus. |
||||
|
Lampanyctus waiohaensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Lampanyctus. |
||||
|
Lashanichthys[65] |
Gen. et comb. nov |
Valid |
Xu et al. |
A member of Ginglymodi assigned to the group Kyphosichthyiformes and to the new family Lashanichthyidae. The type species is "Sangiorgioichthys" sui López-Arbarello et al. (2011); genus also includes "Sangiorgioichthys" yangjuanensis Chen et al. (2014). |
||||
|
Latellopsis[66] |
Gen. et comb. nov. |
Valid |
Bannikov & Zorzin |
Monte Bolca locality |
A member of Perciformes of uncertain phylogenetic placement. The type species is "Psettopsis" latellai Bannikov (2005). |
|||
|
Lepidorhynchus frosti[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Macrouridae. |
||||
|
Leptoscopus atavus[38] |
Sp. nov |
Valid |
Schwarzhans |
|||||
|
Lethrinus? crassiornatus[38] |
Sp. nov |
Valid |
Schwarzhans |
Possibly a species of Lethrinus. |
||||
|
Lindoeichthys[67] |
Gen. et sp. nov. |
Valid |
Murray et al. |
A member of Percopsiformes. Genus includes new species L. albertensis. Announced in 2019; the final version of the article naming it was published in 2020. |
||||
|
Liparis? hampdenensis[38] |
Sp. nov |
Valid |
Schwarzhans |
Possibly a species of Liparis. |
||||
|
Loancorhynchus[68] |
Gen. et sp. nov |
Valid |
Otero |
Middle Eocene |
Millongue Formation |
A relative of the swordfish. The type species is L. catrillancai. |
||
|
Lophiodes hoi[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Lophiodes. |
||||
|
Lophius alveolus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Lophius. |
||||
|
Lophius onokensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Lophius. |
||||
|
Lotella matata[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Lotella. |
||||
|
Lotella pfeili[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Lotella. |
||||
|
Lotella spicata[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Lotella. |
||||
|
Macrurulus depressus[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Merlucciidae. |
||||
|
Macrurulus fordycei[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Merlucciidae. |
||||
|
Maoricottus[38] |
Gen. et comb. et 2 sp. nov |
Valid |
Schwarzhans |
A member of the family Cottidae. The type species is "Cottidarum" impolitus Schwarzhans (1980); genus also includes new species M. calidophilus and M. ovatus. |
||||
|
Melamphaes leeae[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Melamphaes. |
||||
|
Moradebrichthys[69] |
Gen. et sp. nov |
Valid |
Cartanyà et al. |
A member of the family Perleididae. Genus includes new species M. vilasecae. |
||||
|
Muraenanguilla[38] |
Gen. et comb. et 2 sp. nov |
Valid |
Schwarzhans |
A member of Anguilloidei of uncertain phylogenetic placement. The type species is Otolithus (Trachini)" thevenini Priem (1906); genus also includes new species M. balegemensis and M. lacinata. |
||||
|
Myctophum bortonensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Myctophum. |
||||
|
Myctophum tamumuensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Myctophum. |
||||
|
Myctophum tenellum[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Myctophum. |
||||
|
Mystocheilus[39] |
Gen. et sp. nov |
Valid |
Van Tassell & Smith |
A member of the family Cyprinidae. The type species is M. fresti. |
||||
|
Nemadactylus trulliformis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Nemadactylus. |
||||
|
Nemadactylus utoka[38] |
Sp. nov |
Valid |
Schwarzhans |
Otekaike Limestone |
A species of Nemadactylus. |
|||
|
Neobythites turpidus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Neobythites. |
||||
|
Nezumia morgansi[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Nezumia. |
||||
|
Notoconger[38] |
Gen. et 2 sp. nov |
Valid |
Schwarzhans |
A member of the family Congridae. The type species is N. hesperis; genus also includes N. devexus. |
||||
|
Notoscopelus effertus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Notoscopelus. |
||||
|
Notoscopelus praejaponicus[54] |
Sp. nov |
Valid |
Schwarzhans & Ohe |
Chinen Formation |
A species of Notoscopelus. |
|||
|
Opistognathus wharekuriensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Opistognathus. |
||||
|
Oreochromimos[70] |
Gen. et sp. nov |
Penk et al. |
A cichlid belonging to the tribe Oreochromini. The type species is O. kabchorensis. |
|||||
|
Padangia[71] |
Gen. et comb. nov |
Valid |
Murray |
Probably Eocene |
A member of the family Cyprinidae. Genus includes "Sardinioides" amblyostoma von der Marck (1876). |
|||
|
Panturichthys grenfelli[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Panturichthys. |
||||
|
Paralabrus[72] |
Gen. et sp. nov. |
Valid |
Bannikov & Zorzin |
Monte Bolca locality |
Possibly a wrasse. The type species is P. rossiae. |
|||
|
Parapercis bispicatus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Parapercis. |
||||
|
Parapercis depressidorsalis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Parapercis. |
||||
|
Parapercis fatoides[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Parapercis. |
||||
|
Parapercis pareoraensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Parapercis. |
||||
|
Parapercis richardsoni[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Parapercis. |
||||
|
Parapercis waiwaia[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Parapercis. |
||||
|
Parascombrops giganteus[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Acropomatidae. |
||||
|
Parascombrops schwarzhansi[73] |
Sp. nov |
Valid |
Van Hinsbergh & Helwerda |
Late Pliocene to early Pleistocene |
A member of the family Acropomatidae. |
|||
|
Paraulopus rallus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Paraulopus. |
||||
|
Sp. nov |
Wallaard et al. |
A member of Aulopiformes belonging to the family Dercetidae. |
||||||
|
Pempheris hurupiensis[38] |
Sp. nov |
Valid |
Schwarzhans |
Hurupi Formation |
A species of Pempheris. |
|||
|
Phractocephalus yaguaron[75] |
Sp. nov |
Valid |
Bogan & Agnolín |
Late Miocene |
A relative of the redtail catfish. |
|||
|
Physiculus beui[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Physiculus. |
||||
|
Platycephalus? iaiunus[38] |
Sp. nov |
Valid |
Schwarzhans |
Possibly a species of Platycephalus. |
||||
|
Platycephalus? riremoana[38] |
Sp. nov |
Valid |
Schwarzhans |
Possibly a species of Platycephalus. |
||||
|
Sp. nov |
Valid |
Wen et al. |
||||||
|
Protomyctophum ahunga[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Protomyctophum. |
||||
|
Pseudaequalobythites[38] |
Gen. et sp. et comb. nov |
Valid |
Schwarzhans |
A member of the family Ophidiidae. The type species is P. biplex; genus also includes "Otolithus" hilgendorfi Koken (1891). |
||||
|
Pseudanthias multicrenatus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Pseudanthias. |
||||
|
Pseudocaranx? pertenuis[38] |
Sp. nov |
Valid |
Schwarzhans |
Possibly a species of Pseudocaranx. |
||||
|
Pseudophycis muringa[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Pseudophycis. |
||||
|
Pterygotrigla stewarti[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Pterygotrigla. |
||||
|
Sp. nov |
Valid |
Romano et al. |
Possibly a member of the family Turseoidae. |
|||||
|
Sp. nov |
Valid |
Vullo et al. |
||||||
|
Quasinectes[66] |
Gen. et sp. nov. |
Valid |
Bannikov & Zorzin |
Monte Bolca locality |
A member of Perciformes of uncertain phylogenetic placement. The type species is Q. durello. |
|||
|
Rebekkachromis[78] |
Gen. et 2 sp. nov |
Valid |
Kevrekidis, Valtl & Reichenbacher in Kevrekidis et al. |
A cichlid belonging to the subfamily Pseudocrenilabrinae and the tribe Oreochromini.[79] The type species is R. ngororus; genus also includes R. kiptalami. |
||||
|
Rhamphogobius[63] |
Gen. et sp. et comb. nov |
Valid |
Bradić-Milinović, Ahnelt & Schwarzhans in Bradić-Milinović et al. |
Early Miocene |
A member of the family Gobiidae. The type species is R. varidens; genus also includes "Gobius" doppleri Reichenbacher (1993), "Gobius" gregori Reichenbacher (1993) and "Gobius" helvetiae Salis (1967). |
|||
|
Rhynchoconger otaianus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Rhynchoconger. |
||||
|
Saccogaster parengarenga[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Saccogaster. |
||||
|
Sardinella claviformis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Sardinella. |
||||
|
Sardinella lintriculus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Sardinella. |
||||
|
Schernfeldfuro[80] |
Gen. et sp. nov |
Valid |
Ebert |
A member of Halecomorphi. Genus includes new species S. uweelleri. |
||||
|
Scleropages sanshuiensis[81] |
Sp. nov |
Valid |
Zhang |
Early Eocene |
Huachong Formation |
A species of Scleropages. Announced in 2019; the final version of the article naming it was published in 2020. |
||
|
Scopelarchoides neamticus[82] |
Sp. nov |
Valid |
Grădianu et al. |
Lower Dysodilic Shales Formation |
A pearleye, a species of Scopelarchoides. |
|||
|
Scopelosaurus? brevicauda[38] |
Sp. nov |
Valid |
Schwarzhans |
Possibly species of Scopelosaurus. |
||||
|
Sillaginodes albisaxosus[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Sillaginidae. |
||||
|
Sillago maxwelli[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Sillago. |
||||
|
Sprattus arewhana[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Sprattus. |
||||
|
Sundabarbus[71] |
Gen. et comb. nov |
Valid |
Murray |
Probably Eocene |
Sangkarewang Formation |
A member of the family Cyprinidae. Genus includes "Barbus" megacephalus Günther (1876). |
||
|
Symbolophorus moriguchii[54] |
Sp. nov |
Valid |
Schwarzhans & Ohe |
Pliocene (Piacenzian) and Pleistocene (Gelasian) |
Dainichi Formation |
A species of Symbolophorus. |
||
|
Symbolophorus opononiensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Symbolophorus. |
||||
|
Symbolophorus tongaporutuensis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Symbolophorus. |
||||
|
Symphysanodon inamata[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Symphysanodon. |
||||
|
Thalasseleotris whatua[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Thalasseleotris. |
||||
|
Tharsis elleri[83] |
Sp. nov |
Valid |
Arratia, Schultze & Tischlinger |
A teleost belonging to the family Ascalaboidae. |
||||
|
Gen. et comb. nov |
Valid |
Ebert |
A member of the family Pycnodontidae; a new genus for Pycnodus wagneri Thiollière (1852). |
|||||
|
Thryptodus loomisi[85] |
Sp. nov |
Valid |
Shimada |
A member of the family Plethodidae. |
||||
|
Tonganago wharenga[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Congridae. |
||||
|
Tongarewa[38] |
Gen. et 2 sp. nov |
Valid |
Schwarzhans |
A threefin blenny. The type species is T. waihaoensis; genus also includes T. clementsi. |
||||
|
Toroatherina[38] |
Gen. et sp. et comb. nov |
Valid |
Schwarzhans |
An Old World silverside. The type species is T. toroa; genus also includes "Otolithus (Mugilidarum)" debilis Koken (1891). |
||||
|
Toxopyge[63] |
Gen. et sp. et comb. nov |
Valid |
Bradić-Milinović, Ahnelt & Schwarzhans in Bradić-Milinović et al. |
Early Miocene |
A member of the family Gobiidae. The type species is T. campylus; genus also includes "Gobius" longus Salis (1967). |
|||
|
Trachyrincus aulax[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Trachyrincus. |
||||
|
Tranawuen[61] |
Gen. et comb. nov |
Valid |
Gouiric-Cavalli, Remírez & Kriwet |
Early Cretaceous (Valanginian–early Hauterivian) |
A member of Pycnodontiformes. Genus includes "Macromesodon" agrioensis. |
|||
|
Trawdenia[86] |
Gen. et comb. nov |
Valid |
Coates & Tietjen |
An early ray-finned fish. The type species is "Rhadinichthys" planti Traquair (1888); genus also includes "Mesopoma" carricki Coates (1993) and "Mesopoma" pancheni Coates (1993). |
||||
|
Tunisiaclupea[87] |
Gen. et sp. nov |
Valid |
Boukhalfa et al. |
Chotts Basin |
A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Paraclupeidae. Genus includes new species T. speratus. |
|||
|
Uranoscopus rudis[38] |
Sp. nov |
Valid |
Schwarzhans |
Chatton Formation |
A species of Uranoscopus. |
|||
|
Uranoscopus tectiformis[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Uranoscopus. |
||||
|
Valenciennellus fastigatus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Valenciennellus. |
||||
|
Gen. et sp. nov |
Valid |
Díaz-Cruz, Alvarado-Ortega & Carbot-Chanona |
Cintalapa Formation |
A member of Aulopiformes belonging to the family Enchodontidae. Genus includes new species V. avendanoi. |
||||
|
Vinciguerria orientalis[90] |
Sp. nov |
Valid |
Nam, Ko & Nazarkin |
Middle Miocene |
Duho Formation |
A species of Vinciguerria. |
||
|
Waihaoclupea[38] |
Gen. et sp. nov |
Valid |
Schwarzhans |
A member of the family Clupeidae. The type species is W. pinguis. |
||||
|
Waitahana[38] |
Gen. et comb. nov |
Valid |
Schwarzhans |
A southern sandfish. The type species is "Trachinoideorum" sagittiformis Schwarzhans (1980); genus also includes "Citharus" latisulcatus Frost (1924) and "Trachinoideorum" ultimus Schwarzhans (1980). |
||||
|
Waitakia aho[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Hemerocoetidae. |
||||
|
Waitakia proclinens[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Hemerocoetidae. |
||||
|
Waitakia profunda[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Hemerocoetidae. |
||||
|
Xenocephalus otaianus[38] |
Sp. nov |
Valid |
Schwarzhans |
A species of Xenocephalus. |
||||
|
Yudaiichthys[65] |
Gen. et sp. nov |
Valid |
Xu et al. |
A member of Ginglymodi assigned to the group Kyphosichthyiformes and to the new family Lashanichthyidae. The type species is Y. eximius. |
||||
|
Zandtfuro[80] |
Gen. et sp. nov |
Valid |
Ebert |
A member of Halecomorphi. Genus includes new species Z. tischlingeri. |
||||
|
Zonobythites cornifer[38] |
Sp. nov |
Valid |
Schwarzhans |
A member of the family Ophidiidae. |
||||
.svg.png.webp){kind=small}
Lobe-finned fishes
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Anchidipterus[91] |
Gen. et sp. nov |
Valid |
Krupina in Lebedev, Krupina & Linkevich |
A lungfish. Genus includes new species A. dariae. |
||||
|
Clackodus[92] |
Gen. et comb. nov |
Valid |
Smithson, Challands & Smithson |
|||||
|
Sp. nov |
Valid |
Kemp |
Ducabrook Formation |
A lungfish. |
||||
|
Gen. et sp. nov |
Valid |
Gess & Clement |
Witpoort Formation |
A lungfish. The type species is I. mlomomde. |
||||
|
Limanichthys[95] |
Gen. et sp. nov |
Valid |
Challands et al. |
A lungfish. Genus includes new species L. fraseri. |
||||
|
Rubrognathus[96] |
Gen. et sp. nov |
Valid |
Lebedev & Clément |
Yam-Tesovo Formation |
A member of Elpistostegalia. The type species is R. kuleshovi. |
|||
|
Selenodus[97] |
Gen. et sp. nov |
Valid |
Mondéjar-Fernández |
An onychodont sarcopterygian. Genus includes new species S. aquesbiae. |
||||
|
Sp. nov |
Valid |
Yabumoto et al. |
Middle Sakamena Group |
|||||
.svg.png.webp){kind=small}
Other fishes
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Aberrosquama[99] |
Gen. et sp. nov |
Valid |
Burrow in Burrow et al. |
Hamelin Formation |
A stem-gnathostome of uncertain phylogenetic placement. The type species is A. occidens. |
|||
General research
- Revision of Early Devonian psammosteids described by Beverly Halstead from the so-called "Placoderm Sandstone" (Świętokrzyskie Mountains, Poland) is published by Dec (2019).[100]
- A study on the locomotion of psammosteids, focusing on the influence of the shape of the psammosteid body form on hydrodynamic performance, is published by Dec (2019).[101]
- A study on the anatomy of dermal plates of Astraspis is published by Lemierre & Germain (2019), who report possible evidence of presence of proliferative cartilage in this taxon.[102]
- A study on the anatomy of the circulatory system of the head of Shuyu is published by Gai, Zhu & Donoghue (2019).[103]
- Redescription of Sinogaleaspis shankouensis, based on data from 11 new specimens from the Silurian Xikeng Formation (Jiangxi, China), is published online by Gai et al. (2019).[104]
- A study on the anatomy of the dermal skeleton of Tremataspis mammillata is published by O'Shea, Keating & Donoghue (2019).[105]
- Redescription of Asterolepis orcadensis based on newly collected fossil material is published by Newman, den Blaauwen & Leather (2019).[106]
- A study on the morphology of the jaw elements of a buchanosteoid placoderm specimen ANU V244 from the Early Devonian limestones (~400 Ma) at Burrinjuck, near Canberra (Australia) is published by Hu et al. (2019).[107]
- A specimen of Dunkleosteus terrelli preserving vertebrae fused into a structure known as the synarcual is described from the Devonian Cleveland Shale Member of the Ohio Shale Formation (Ohio, United States) by Johanson et al. (2019).[108]
- A redescription and a study on the phylogenetic relationships of a putative antiarch Silurolepis platydorsalis is published by Zhu, Lu & Zhu (2019), who reinterpret this species as a maxillate placoderm close to Qilinyu.[109]
- New body fossils of Cheiracanthus intricatus, including the first known articulated specimen of this species, are described from the Givetian Tordalen Formation (Spitsbergen, Norway) by Newman, Burrow & den Blaauwen (2019).[110]
- A study on the anatomy of the skeletal elements of the pharynx of Ptomacanthus anglicus, and on its implications for the knowledge of the evolution of the pharynx of jawed vertebrates, is published by Dearden, Stockey & Brazeau (2019).[111]
- Description of fossils of Carboniferous (Mississippian) cartilaginous fishes from the area of Krzeszowice (Poland) is published by Ginter & Złotnik (2019).[112]
- A study on microwear on teeth of Edestus minor, and on its implications for the knowledge of function of teeth of this fish, is published by Itano (2019).[113]
- A revision of species belonging to the genus Edestus is published by Tapanila & Pruitt (2019).[114]
- Discovery of a cast of the holotype of Petalodus ohioensis in the collections of the Yale Peabody Museum of Natural History is reported by Carpenter & Itano (2019), who consider the species Petalodus alleghaniensis to be a junior synonym of P. ohioensis.[115]
- Restudy of a putative bill of an ibis-like bird from the Eocene La Meseta Formation (Antarctica) described by Jadwiszczak, Gaździcki & Tatur (2008)[116] is published by Agnolin, Bogan & Rozadilla (2019), who consider this specimen to be more likely to be a dorsal spine of a chimaeroid cartilaginous fish.[117]
- A study on the anatomy of the skull of Tristychius arcuatus, providing evidence of adaptations for suction feeding, is published by Coates et al. (2019).[118]
- Description of the first skeletal remains of Phoebodus from the Famennian of the Maïder region of Morocco, providing new information on the anatomy of this species, and a study on the phylogenetic affinities of Phoebodus is published by Frey et al. (2019).[119]
- Teeth of a hybodont shark belonging to the genus Asteracanthus, with anatomy indicative of a crushing feeding behaviour, are described from the Upper Jurassic deposits of the Monte Nerone Pelagic Carbonate Platform, in the Umbria‐Marche‐Sabina Palaeogeographic Domain (Italy) by Citton et al. (2019).[120]
- An association of 58 teeth of Ptychodus anonymus, representing the first occurrence of an associated dentition of this species, is described from the Cenomanian Jetmore Member of the Greenhorn Formation (Kansas, United States) by Hamm (2019).[121]
- Description of new associated skeletal remains of Ischyrhiza mira from the Upper Cretaceous of Tennessee and Alabama and a study on the paleobiology of this species is published by Sternes & Shimada (2019).[122]
- A study on the anatomy and phylogenetic relationships of Promyliobatis gazolai is published by Marramà et al. (2019).[123]
- Fossils of members of the genus Aetomylaeus are described from localities in Peru and Chile by Villafaña et al. (2019), representing the first unambiguous fossil record of this genus from the Neogene of the southeastern Pacific.[124]
- A specimen of the whiptail stingray species Tethytrygon muricatus preserving the uterus bearing four eggs is described from the Eocene of the Monte Bolca locality (Italy) by Fanti, Mazzuferi & Marramà (2019).[125]
- Description of Langhian ray fossils from the Lower Tagus Basin (Portugal) is published by Fialho, Balbino & Antues (2019).[126]
- A study on the taxonomic status and geological age of large shark remains from the Upper Cretaceous of the Castellavazzo locality (Italy) discovered in the 19th century is published by Conte et al. (2019).[127]
- 15 partial skeletons of lamniform sharks, including the largest specimen of Cretoxyrhina mantelli known to date, are described from the Upper Cretaceous Scaglia Rossa Formation (Italy) by Amalfitano et al. (2019), who also review the taxonomic history of C. mantelli.[128]
- A study on teeth histology and mineralization pattern in lamniform sharks, based on data from extant and fossil taxa (including enigmatic galeomorph shark Palaeocarcharias stromeri), is published by Jambura et al. (2019).[129]
- A study on changes of diversity of lamniform sharks throughout their evolutionary history, aiming to determine the causes of their decline in the last 20 million years, is published by Condamine, Romieu & Guinot (2019).[130]
- A study on changes in the presence or absence of lateral cusplets on teeth of members of the genus Carcharocles from the Calvert, Choptank, and St. Marys formations, and on their implications for the transition of shark populations from these formations from a Carcharocles chubutensis-dominated population to one dominated by C. megalodon, is published by Perez et al. (2019).[131]
- A revision and a reevaluation of the reliability of all post-Messinian occurrences of Otodus megalodon in marine strata from western North America, and a study on the timing of extinction of this species, is published by Boessenecker et al. (2019).[132]
- A study on the body size of Otodus megalodon, as inferred from the relationship between the ontogenetic development of teeth and total body length in the great white shark, is published by Shimada (2019).[133]
- Partial forelimb of a rorqual with several shark bite marks is described from the Pliocene Burica Formation (Panama) by Cortés et al. (2019).[134]
- A study on the anatomy of the holotype specimen of a putative Paleocene shark Platyacrodus unicus is published by Bogan, Agnolin & Ezcurra (2019), who reinterpret this specimen as a carapace of a small retroplumid crab belonging to the genus Costacopluma.[135]
- An assemblage of well-preserved isolated teeth of elasmobranchs is described from the late Oligocene of the North Alpine Foreland Basin (Austria) by Feichtinger et al. (2019).[136]
- Description of the deep-sea elasmobranch fauna from the Miocene Yatsuo Group in central Japan, including the first fossil occurrences of the genera Arhynchobatis and Pseudoraja, will be published by Nishimatsu & Ujihara (2019).[137]
- Description of the deep-sea elasmobranch fossils from the Miocene Makino Formation in southwest Japan, including the first fossil occurrences of the genera Springeria and Narke, is published by Nishimatsu (2019).[138]
- A study on Paleocene cartilaginous fish fossils from the Lower Clayton Limestone Unit of the Midway Group near Malvern, Arkansas, evaluating the implications of these fossils for the knowledge of cartilaginous fish diversity across the Cretaceous-Paleogene boundary in the Malvern region and Gulf Coastal Plain of southwestern Arkansas, is published online by Maisch, Becker & Griffiths (2019).[139]
- A study on the morphology of scales and squamation pattern in Guiyu oneiros is published by Cui, Qiao & Zhu (2019).[140]
- A study on the anatomy and phylogenetic relationships of Brazilichthys macrognathus is published by Figueroa, Friedman & Gallo (2019).[141]
- Permian species Palaeothrissum inaequilobum Blainville (1818) and P. parvum Blainville (1818) are found to be senior synonyms of the widely used species name Aeduella blainvillei (Agassiz, 1833) by Brignon (2019). Conditions exist for reversal of precedence and Aeduella blainvillei is declared nomen protectum.[142]
- A study on the anatomy of the jaws, palate and teeth of Eurynotus crenatus is published by Friedman et al. (2019).[143]
- New specimen of Birgeria liui, representing the most complete and articulated postcranial skeleton of an adult specimen of Birgeria reported so far, is described from the Ladinian of South China by Ni et al. (2019).[144]
- Fossil remains of a spiral valve are reported in a specimen of Peipiaosteus pani from the Lower Cretaceous Yixian Formation (China) by Capasso (2019).[145]
- Teeth of members of the genus Pycnodus are described from the Upper Cretaceous Cap de Naze marine formation (Senegal) by Capasso (2019).[146]
- Description of a partial skeleton of Micropycnodon kansasensis from the Smoky Hill Chalk Member of the Niobrara Chalk (Kansas, United States), providing new information on the anatomy of this species, is published by Cronin & Shimada (2019).[147]
- A study on the paleobiology of specimens of Nursallia gutturosum from the Cenomanian-Turonian platy limestone deposit of Vallecillo (north-eastern Mexico) is published by Stinnesbeck, Rust & Herder (2019).[148]
- A study on the anatomy and phylogenetic relationships of Robustichthys luopingensis is published by Xu (2019).[149]
- Fossil teeth representing the first evidence of Late Jurassic ginglymodians from Mt. Nerone in the Umbria-Marche-Sabina Domain (Italy) are described by Romano et al. (2019).[150]
- A study on the anatomy and phylogenetic relationships of "Lepidotes" bernissartensis is published online by Cavin, Deesri & Olive (2019), who transfer this species to the genus Scheenstia.[151]
- A study on the internal anatomy of a left jaw of a member of the genus Scheenstia from the Kimmeridgian Reuchenette Formation (Switzerland) is published online by Leuzinger et al. (2019), who describe a peculiar tooth replacement mode in this specimen.[152]
- A study on the stomach contents of two specimens of Lepidotes from the Lower Jurassic of Germany is published online by Thies, Stevens & Stumpf (2019).[153]
- Six new occurrences of Belonostomus, documenting the biogeographic and biostratigraphic range of this genus in North America, are reported from the Upper Cretaceous of Texas, Alabama and Mississippi by Van Vranken, Fielitz & Ebersole (2019).[154]
- A study on the diversity of pectoral fin shape amongst members of Pachycormiformes is published by Liston et al. (2019).[155]
- A study on the anatomy of the skull of Martillichthys renwickae is published online by Dobson et al. (2019).[156]
- Fossil remains of a member or a relative of the genus Asthenocormus are described from the Upper Jurassic of the Ameghino (= Nordenskjöld) Formation of the Antarctic Peninsula by Gouiric-Cavalli et al. (2019), representing the first record of a suspension-feeding pachycormid from the Upper Jurassic of the Antarctic Peninsula and the oldest pachycormid yet recovered from Antarctica.[157]
- New, three-dimensionally preserved specimens of Pachycormus are described from the Toarcian of Strawberry Bank at Ilminster (Somerset, United Kingdom) by Cawley et al. (2019).[158]
- A study on the anatomy of Pleuropholis decastroi from the Lower Cretaceous (Albian) limestones of Pietraroja (Province of Benevento, Italy) and on the phylogenetic relationships of the family Pleuropholidae is published by Taverne & Capasso (2019).[159]
- A study on the anatomy and phylogenetic relationships of the teleost species Majokia brasseuri from the Middle Jurassic Stanleyville Formation (Democratic Republic of the Congo) is published by Taverne (2019), who names a new order Majokiiformes.[160]
- A study on the anatomy, jaw mechanics and phylogenetic relationships of Dugaldia emmilta is published by Cavin & Berrell (2019).[161]
- A juvenile specimen of Xiphactinus audax, representing the smallest specimen of this species reported so far, will be described from the Cretaceous Niobrara Chalk (Kansas, United States) by King & Super (2019).[162]
- A study on the internal structure of a fossil specimen of Notelops brama, investigated by neutron tomography, is published by Pugliesi et al. (2019).[163]
- A study on the anatomy and phylogenetic relationships of Cavenderichthys talbragarensis and Waldmanichthys koonwarri is published online by Bean & Arratia (2019).[164]
- A study on fossils of members of the genus Capoeta from the Pliocene locality Çevirme (Turkey) and on the evolutionary history of this genus is published by Ayvazyan, Vasilyan & Böhme (2019).[165]
- Evidence of presence of managed aquaculture of the common carp by around 6000 BC is reported from the Early Neolithic Jiahu site (China) by Nakajima et al. (2019).[166]
- A study on the morphology of fossil catfish spines from the Upper Cretaceous Adamantina and Marilia formations (Brazil) is published by Alves, Bergqvist & Brito (2019).[167]
- A study on positions, heading directions and possible behavioural rules used in a group of fossil specimens of Erismatopterus levatus from the Eocene Green River Formation is published by Mizumoto, Miyata & Pratt (2019).[168]
- Description of gadiform fossils from the Eocene sediments of the Sverdlovsk and Tyumen regions (Ural and Western Siberia, Russia), including fossils of members of the family Merlucciidae, is published by Marramà et al. (2019).[169]
- Description of an incomplete percomorph specimen from the Miocene Ixtapa Formation (Mexico), representing the oldest primary freshwater percomorph fish from Mexico reported so far, is published by Cantalice & Alvarado-Ortega (2019).[170]
- A study on the evolutionary history of the fish clade Pelagiaria is published by Friedman et al. (2019).[171]
- A revision of the nomenclature of extant and fossil barracudas is published by Ballen (2019).[172]
- A review of the published fossil record of the family Labridae is published by Bellwood et al. (2019).[173]
- A study on the feeding habits of the percomorph fish Rhenanoperca minuta and other fishes from the Eocene Messel pit (Germany) is published by Micklich, Baranov & Wappler (2019).[174]
- A study on the phylogenetic relationships of "psarolepid" bony fishes, evaluating which characters cause their different placements in analyses utilizing different methods for reconstructing the tree of life, is published by King (2019).[175]
- A study on the ontogeny of the neurocranium and brain in the West Indian Ocean coelacanth, and on its implications for the knowledge of the evolution of the head of lobe-finned fishes, is published by Dutel et al. (2019).[176]
- Redescription of Axelrodichthys araripensis and a comparative study of several other members of Mawsoniidae is published by Fragoso, Brito & Yabumoto (2019), who transfer the species Mawsonia lavocati to the genus Axelrodichthys.[177]
- A study on the phylogenetic relationships and evolutionary history of mawsoniid coelacanths is published by Cavin et al. (2019).[178]
- A study on the skull anatomy of Arquatichthys porosus, focusing on a newly-discovered postparietal shield, is published by Lu & Zhu (2019).[179]
- Description of the posterior part of the skull of Tungsenia paradoxa is published by Lu et al. (2019).[180]
- New fossil material of "Holoptychius" radiatus Newberry (1889) from the Devonian (Famennian) Catskill Formation (Pennsylvania, United States), providing new information on the anatomy of this species, is described by Daeschler, Downs & Matzko (2019), who transfer this species to the tristichopterid genus Langlieria.[181]
- A study on the bone histology of the humerus of Hyneria lindae is published by Kamska et al. (2019).[182]
- Description of new fossil material of Edenopteron from the Devonian (Famennian) Worange Point Formation (Australia) and a study on the phylogenetic relationships of this taxon is published online by Young et al. (2019).[183]
- A study on the anatomy of the shoulder girdle and opercular series of Gogonasus andrewsae is published by Hu, Young & Lu (2019).[184]
- A historical review of the fossil record of Devonian tetrapods and basal tetrapodomorphs from East Gondwana (Australasia, Antarctica) is published by Long, Clement & Choo (2019), who also present preliminary findings on the anatomy of the canowindrid Koharalepis jarviki based on synchrotron scan data.[185]
- A study on the anatomy of fossil coelacanth lungs, on accessory air-breathing structures in fossil fishes and stem-tetrapods, and on the evolution of air breathing is published by Cupello, Clément & Brito (2019).[186]
- A study on patterns of tooth replacement in Onychodus jandemarrai, Eusthenopteron foordi, Tiktaalik roseae and in extant West Indian Ocean coelacanth is published by Doeland et al. (2019).[187]
- A study on changes of the skeletal anatomy of the pelvic and pectoral appendages during the transition from fins to limbs in vertebrate evolution, as indicated by data from fossil lobe-finned fishes and early tetrapods, is published by Esteve-Altava et al. (2019).[188]
- A study on the anatomy of dermal rays in pectoral fins of Sauripterus taylori, Eusthenopteron foordi and Tiktaalik roseae, evaluating its implications for the knowledge of the evolution of dermal rays in early members of Tetrapodomorpha prior to the origin of digits, is published online by Stewart et al. (2019).[189]
- A study on the evolution of the branchiostegal ray series in the skull of bony fishes, as indicated by data from extant and fossil taxa, is published by Ascarrunz et al. (2019).[190]
- A study on the origin and evolution of acellular bone (bone without osteocytes) in fossil and extant actinopterygian fishes is published by Davesne et al. (2019).[191]
- A diverse fish assemblage is reported from the Carboniferous (upper Pennsylvanian) Horquilla Formation (New Mexico, United States) by Ivanov & Lucas (2019).[192]
- A study on the diversity and ecology of Triassic fish assemblages from the Villány Hills (Hungary) is published by Szabó, Botfalvai & Osi (2019).[193]
- Cretaceous (probably Barremian–Aptian) fish fossils with strong affinities with Early Cretaceous faunas of Thailand are described from Pahang (Malaysia) by Teng et al. (2019).[194]
- Description of a freshwater fish assemblage from the Cretaceous (Albian-Cenomanian) Açu Formation (Brazil) is published by Veiga, Bergqvist & Brito (2019).[195]
- Description of Eocene (Bartonian) fish fauna from the Luna de Sus locality (Romania) is published by Trif, Codrea & Arghiuș (2019).[196]
- A study on the composition of the otolith assemblage from the Santa Barbara Basin near the coast of California over the preceding two millennia is published by Jones & Checkley (2019).[197]
- A study on the evolution of the herbivorous coral reef fishes, as indicated by data from extant and fossil species, is published by Siqueira, Bellwood & Cowman (2019).[198]
References
- 1 2 3 Tiiu Märss (2019). "Silurian cyathaspidid heterostracans of Northern Eurasia". Estonian Journal of Earth Sciences. 68 (3): 113–146. doi:10.3176/earth.2019.11. S2CID 202181325.
- 1 2 Yu-Hai Liu; Min Zhu; Xiang-Hong Lin; Li-Wu Lu; Zhi-Kun Gai (2019). "A reappraisal of the Silurian galeaspids (stem-Gnathostomata) from Tarim Basin, Xinjiang". Vertebrata PalAsiatica. 57 (4): 253–273. doi:10.19615/j.cnki.1000-3118.190524.
- 1 2 3 Xin-Dong Cui; Qiang Li; Tuo Qiao; Min Zhu (2020). "New material of thelodonts from Lochkovian (Lower Devonian) of Qujing, Yunnan, China". Vertebrata PalAsiatica. 58 (1): 1–15. doi:10.19615/j.cnki.1000-3118.190612.
- ↑ Tetsuto Miyashita; Michael I. Coates; Robert Farrar; Peter Larson; Phillip L. Manning; Roy A. Wogelius; Nicholas P. Edwards; Jennifer Anné; Uwe Bergmann; A. Richard Palmer; Philip J. Currie (2019). "Hagfish from the Cretaceous Tethys Sea and a reconciliation of the morphological–molecular conflict in early vertebrate phylogeny". Proceedings of the National Academy of Sciences of the United States of America. 116 (6): 2146–2151. Bibcode:2019PNAS..116.2146M. doi:10.1073/pnas.1814794116. PMC 6369785. PMID 30670644.
- ↑ Xian-Ren Shan; Xiang-Hong Lin; Yu-Meng Zhang; Xu-Tong Li; Zhi-Kun Gai (2023). "New findings of Xiyuichthys (Xiushuiaspidae, Galeaspida) from the Silurian of Jiangxi Province and Tarim Basin". Vertebrata PalAsiatica. 61 (4): 245–260. doi:10.19615/j.cnki.2096-9899.230904.
- ↑ Jason P. Downs; Edward B. Daeschler; Nathanael Lo; Emily N. Carey; Neil H. Shubin (2019). "Asterolepis alticristata n. sp. (Antiarchi) from the Upper Devonian (Frasnian) of Nunavut, Canada, and a report on the antiarch diversity of the Fram Formation". Geodiversitas. 41 (19): 679–698. doi:10.5252/geodiversitas2019v41a19. S2CID 210624573.
- 1 2 Sébastien Olive; Alan Pradel; Carlos Martinez-Pérez; Philippe Janvier; James C. Lamsdell; Pierre Gueriau; Nicolas Rabet; Philippe Duranleau-Gagnon; Andres L. Cardenas-Rozo; Paula A. Zapata Ramirez; Héctor Botella (2019). "New insights into Late Devonian vertebrates and associated fauna from the Cuche Formation (Floresta Massif, Colombia)". Journal of Vertebrate Paleontology. 39 (3): e1620247. Bibcode:2019JVPal..39E0247O. doi:10.1080/02724634.2019.1620247. hdl:10784/26749. S2CID 198237241.
- ↑ Kate Trinajstic; John A. Long; Alexander O. Ivanov; Elga Mark-Kurik (2019). "A new genus of ptyctodont (Placodermi) from the Late Devonian of Baltic area". Palaeontologia Electronica. 22 (2): Article number 22.2.23. doi:10.26879/890. hdl:1885/222039. S2CID 181769139.
- ↑ Jan L. den Blaauwen; Michael J. Newman; Carole J. Burrow (2019). "A new cheiracanthid acanthodian from the Middle Devonian (Givetian) Orcadian Basin of Scotland and its biostratigraphic and biogeographical significance". Scottish Journal of Geology. 55 (2): 166–177. Bibcode:2019ScJG...55..166D. doi:10.1144/sjg2018-023. S2CID 189990496.
- ↑ Michael J. Newman; Carole J. Burrow; Jan L. den Blaauwen (2019). "A new species of ischnacanthiform acanthodian from the Givetian of Mimerdalen, Svalbard". Norwegian Journal of Geology. 99 (4): 619–631. doi:10.17850/njg99-4-05. S2CID 214220714.
- ↑ Sebastian Stumpf; Jürgen Kriwet (2019). "A new Pliensbachian elasmobranch (Vertebrata, Chondrichthyes) assemblage from Europe, and its contribution to the understanding of late Early Jurassic elasmobranch diversity and distributional patterns". PalZ. 93 (4): 637–658. doi:10.1007/s12542-019-00451-4. S2CID 181782998.
- ↑ Eduardo Villalobos-Segura; Charlie J. Underwood; David J. Ward; Kerin M. Claeson (2019). "The first three-dimensional fossils of Cretaceous sclerorhynchid sawfish: Asflapristis cristadentis gen. et sp. nov., and implications for the phylogenetic relations of the Sclerorhynchoidei (Chondrichthyes)" (PDF). Journal of Systematic Palaeontology. 17 (21): 1847–1870. doi:10.1080/14772019.2019.1578832. S2CID 145940997.
- ↑ Gerard R. Case; Todd D. Cook; Taylor Kightlinger; Paul D. Borodin (2019). "Middle Campanian euselachian diversity of the southern region of the Atlantic Coastal Plain of North America". Vertebrate Anatomy Morphology Palaeontology. 7: 69–82. doi:10.18435/vamp29345. S2CID 201576213.
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- ↑ Karen E. Samonds; Tsiory H. Andrianavalona; Lane A. Wallett; Iyad S. Zalmout; David J. Ward (2019). "A middle - late Eocene neoselachian assemblage from nearshore marine deposits, Mahajanga Basin, northwestern Madagascar". PLOS ONE. 14 (2): e0211789. Bibcode:2019PLoSO..1411789S. doi:10.1371/journal.pone.0211789. PMC 6392239. PMID 30811516.
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