Pachyosteomorphi
Temporal range: Devonian
Artist's reconstruction of various Cleveland Shale Pachyosteomorphi arthrodires, including Titanichthys (top), Selenosteus (eating a shark), adult and juvenile Dunkleosteus terrelli (center), and Gorgonichthys (bottom). From the Famennian stage of the Late Devonian.
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Placodermi
Order: Arthrodira
Suborder: Brachythoraci
Clade: Eubrachythoraci
Clade: Pachyosteomorphi
Stensiö, 1963
Sub-clades

Pachyosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.[1]

Phylogeny

Pachyosteomorphi is the sister taxon to Coccosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Pachyosteomorphi can be further sub-divided into Aspinothoracidi and Dunkleosteoidea,[2][1] as shown in the cladogram below:[3]

Eubrachythoraci
Coccosteomorphi
Coccosteoidea
Coccosteidae

Millerosteus minor

Coccosteus cuspidatus

Dickosteus threiplandi

Watsonosteus fletti

Protitanichthys rockportensis

Panxiosteidae

Plourdosteus canadensis

Panxiosteus ocullus

Janiosteus timanicus

Incisoscutoidea

Harrytoombsia elegans

Torosteus tuberculatus

Torosteus pulchellus

Mcnamaraspis kaprios

Compagopiscis croucheri

Trematosteus fontanellus

Camuropiscidae

Incisoscutum ritchiei

Incisoscutum sarahae

Rolfosteus canningensis

Tubonasus lennardensis

Fallacosteus turneri

Camuropiscis laidlawi

Latocamurus coulthardi

Pachyosteomorphi

Rhachiosteus pterygiatus

Dunkleosteoidea

Eastmanosteus calliaspis

Eastmanosteus pustulosus

Kiangyousteus yohii

Golshanichthys asiatica

Westralichthys uwagedensis

Dunkleosteus raveri

Dunkleosteus terrelli

Dunkleosteus amblyodoratus

Heterostiidae

Heterosteus ingens

Yinosteus major

Aspinothoracidi

Tapinosteus heintzi

Bullerichthys fascidens

Kendrickichthys cavernosus

Bruntonichthys multidens

Dinichthys herzeri

Hadrosteus rapax

Gorgonichthys clarki

Selenosteidae

Heintzichthys gouldii

Pachyosteus bulla

Gymnotrachelus hydei

Stenosteus angustopectus

Brachyosteus dietrichi

Melanosteus occitanus

Rhinosteus parvulus

References

  1. 1 2 You-An Zhu; Min Zhu (2013). "A redescription of Kiangyousteus yohii (Arthrodira: Eubrachythoraci) from the Middle Devonian of China, with remarks on the systematics of the Eubrachythoraci". Zoological Journal of the Linnean Society. 169 (4): 798–819. doi:10.1111/zoj12089.
  2. Carr, R.K.; Hlavin, W.J. (2010). "Two new species of Dunkleosteus Lehman, 1956, from the Ohio Shale Formation (USA, Famennian) and the Kettle Point Formation (Canada, Upper Devonian), and a cladistic analysis of the Eubrachythoraci (Placodermi, Arthrodira)". Zoological Journal of the Linnean Society. 159 (1): 195–222. doi:10.1111/j.1096-3642.2009.00578.x.
  3. Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society. 176 (4): 806–834. doi:10.1111/zoj.12356. ISSN 0024-4082.
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