Coccosteidae
Temporal range: Late Emsian-Late Devonian,
Artist's reconstruction of Coccosteus cuspidatus
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Placodermi
Order: Arthrodira
Suborder: Brachythoraci
Clade: Eubrachythoraci
Clade: Coccosteomorphi
Superfamily: Coccosteoidea
Family: Coccosteidae
Traquir, 1888
Genera

Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China.

Phylogeny

Coccosteidae belongs to the larger clade Coccosteomorphi, which together with its sister clade Pachyosteomorphi forms the group Eubrachythoraci. The phylogeny of Coccosteidae can be shown in the cladogram below:[1]

Eubrachythoraci
Coccosteomorphi
Coccosteoidea
Coccosteidae

Millerosteus minor

Coccosteus cuspidatus

Dickosteus threiplandi

Watsonosteus fletti

Protitanichthys rockportensis

Panxiosteidae

Plourdosteus canadensis

Panxiosteus ocullus

Janiosteus timanicus

Incisoscutoidea

Harrytoombsia elegans

Torosteus tuberculatus

Torosteus pulchellus

Mcnamaraspis kaprios

Compagopiscis croucheri

Trematosteus fontanellus

Camuropiscidae

Incisoscutum ritchiei

Incisoscutum sarahae

Rolfosteus canningensis

Tubonasus lennardensis

Fallacosteus turneri

Camuropiscis laidlawi

Latocamurus coulthardi

Pachyosteomorphi

Rhachiosteus pterygiatus

Dunkleosteoidea

Eastmanosteus calliaspis

Eastmanosteus pustulosus

Kiangyousteus yohii

Golshanichthys asiatica

Westralichthys uwagedensis

Dunkleosteus raveri

Dunkleosteus terrelli

Dunkleosteus amblyodoratus

Heterostiidae

Heterosteus ingens

Yinosteus major

Aspinothoracidi

Tapinosteus heintzi

Bullerichthys fascidens

Kendrickichthys cavernosus

Bruntonichthys multidens

Dinichthys herzeri

Hadrosteus rapax

Gorgonichthys clarki

Selenosteidae

Heintzichthys gouldii

Pachyosteus bulla

Gymnotrachelus hydei

Stenosteus angustopectus

Brachyosteus dietrichi

Melanosteus occitanus

Rhinosteus parvulus

Genera

Belgiosteus

A genus of very large coccosteids. Species are found in Middle Devonian Belgium and China.

Clarkosteus

Coccosteus

The type genus of the family. Numerous species are found in Middle to Upper Devonian strata throughout Europe and parts of North America.

Dickosteus

Jiuchengia

The earliest known coccosteid from Late Emsian Yunnan province, China. It is distinguished from other coccosteids by having an elongated occipital.

Livosteus

A genus of very large coccosteids known from Middle to Late Devonian strata of Eastern Europe.

Millerosteus

A genus of very small coccosteids known from Middle to Late Devonian strata throughout Europe. Some species are found in the same strata with certain species of Coccosteus.

Protitanichthys

Watsonosteus

Woodwardosteus

References

  1. Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society. 176 (4): 806–834. doi:10.1111/zoj.12356. ISSN 0024-4082.
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