Hemipenthes
Hemipenthes maura
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Family: Bombyliidae
Subfamily: Anthracinae
Tribe: Villini
Genus: Hemipenthes
Loew, 1869
Type species
Musca morio
Synonyms

Hemipenthes is a large genus of flies belonging to the family Bombyliidae (bee-flies). There are many described species, distributed throughout the Holarctic realm. These are small to large robust flies with a body length of 5–14 mm. They can be distinguished from similar genera (Villa) by their wing venation.[1] A number of species formerly in this genus were moved to a separate genus, ins in 2020.[2]

Larvae are hyperparasites on parasitic Hymenoptera.[1]

Extant species

  • H. alba Ávalos-Hernández, 2009 - Nearctic: Mexico[3]
  • H. bigradata (Loew, 1869) - Nearctic: USA (California, New Mexico). Neotropical: Cuba, Bahamas, Jamaica, Puerto Rico
  • H. blanchardiana (Jaennicke, 1867) - Nearctic: Mexico (Distrito Federal, Guanajuato, Guerrero, México, Puebla, Sonora), USA (Arizona, California, Texas)
  • H. castanipes Bigot, 1892 - Nearctic
  • H. catulina (Coquillett, 1894) - Nearctic: USA (California, Colorado, Idaho, Illinois, Indiana, Iowa, Michigan, Minnesota, Montana, New Jersey, New Mexico, New York, Ohio, Pennsylvania, Utah, Washington, Wisconsin, Wyoming)
  • H. chimaera (Osten Sacken, 1887) - Nearctic: Mexico (Guerrero, Sonora), USA (Arizona)
  • H. comanche (Painter in Painter & Painter, 1962) - Nearctic: USA (Arizona, California, Colorado, Nevada, New Mexico, Utah)
  • H. differens (Hall, 1976) - Neotropical: Chile
  • H. discolor (Hall, 1976) - Neotropical: Chile
  • H. ditaenia (Wiedemann, 1828) - Neotropical: Argentina, Chile, Peru, Uruguay
  • H. divisa (Walker) - Neotropical: South America
  • H. edwardsii (Coquillett, 1894) - Nearctic: Canada (British Columbia), USA (California)
  • H. edwarsi (Oldroyd, 1938) - Neotropical: Chile
  • H. epilais (Wiedemann, 1828) - Neotropical: Brazil
  • H. ethiops (Greathead, 1967) - Afrotropical: Ethiopia
  • H. eversmanni Zaitzev and Madra Mandicencio, 1966 - Palaearctic: Kazakhstan, Moldova, Russia, Ukraine
  • H. exoprosopoides Paramonov, 1928 - Palaearctic: Armenia, Azerbaijan, China (Sichuan), Iran, Kyrgyz Republic, Tajikistan, Turkmenistan, Uzbekistan.
  • H. extensa Wulp, 1888 - Neotropical: Argentina
  • H. galapagensis (Painter & Painter, 1974) - Neotropical: Argentina
  • H. galathea (Osten Sacken, 1886) - Neotropical: Costa Rica
  • H. gaudanica Paramonov, 1927 - Palaearctic: China (Xinjiang), Iran, Kyrgyz Republic, Mongolia, Tajikistan, Turkmenistan, Uzbekistan
  • H. gayi (Macquart, 1840) - Neotropical: Chile
  • H. gussakovskyi Zaitzev, 1966 - Palaearctic: Tajikistan, Turkmenistan
  • H. hamifera (Loew, 1854) - Palaearctic: Armenia, Azerbaijan, Bulgaria, China (Jiangsu, Nei Monggol, Xinjiang), France, Greece, Gruzia, Iran, Italy (incl. Sicily), Kazakhstan, Kyrgyz Republic, Mongolia, Russia (ES, WS), Spain, Tajikistan, Turkey, Turkmenistan, Uzbekistan, Yugoslavia
  • H. incisiva (Walker, 1852) - Nearctic: Mexico (Guerrero)
  • H. inops (Coquillett, 1887) - Nearctic: USA (Arizona, California, Colorado, Oregon, Utah, Washington)
  • H. jaennickeana (Osten Sacken, 1886) - Nearctic: Mexico (Morelos, Sonora), USA (Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, Texas, Utah)
  • H. jezoensis (Matsumura, 1916) - Oriental: Taiwan. Palaearctic: Japan
  • H. lepidota (Osten Sacken, 1887) - Nearctic: Canada (Alberta), Mexico (Baja California Norte, Baja California Sur, Chihuahua, Guerrero, Puebla, San Luis Potosí, Sonora, Tamaulipas), USA (Arizona, California, Colorado, Idaho, Louisiana, Nevada)
  • H. maura (Linnaeus, 1758) - Palaearctic
  • H. melaleuca (Wiedemann, 1828) - Neotropical: Argentina, Uruguay
  • H. melana Bowden, 1965 - Oriental: Nepal
  • H. mesasiatica Zaitzev, 1962 - Palaearctic: Tajikistan, Turkmenistan, Uzbekistan
  • H. micromelas (Bigot, 1892) - Neotropical: Argentina
  • H. mischanensis Paramonov, 1927 - Palaearctic: Armenia
  • H. montanorum (Austen, 1936) - Oriental: China (Yunnan). Palaearctic: China (Qinghai, Sichuan, Xizang)
  • H. morio (Linnaeus, 1758) - Oriental, Palaearctic
  • H. nitidofasciata (Portschinsky, 1892) - Palaearctic: Kyrgyz Republic, Russia, Tajikistan
  • H. noscibilis (Austen, 1936) - Palaearctic: China (Xizang)
  • H. nudiuscula (Thomson, 1869) - Neotropical: Panama
  • H. pamirensis Zaitzev, 1962 - Palaearctic: Mongolia, Tajikistan, Turkmenistan, Uzbekistan
  • H. panfilovi Zaitzev, 1981 - Palaearctic: Russia
  • H. pauper (Becker, 1916) - Palaearctic: Algeria, Egypt
  • H. praecisa (Loew, 1869) - Palaearctic: China (Beijing, Hebei, Nei Monggol), Kyrgyz Republic, Mongolia, Russia, Tajikistan, Turkmenistan, Uzbekistan
  • H. pullata (Coquillett, 1894) - Nearctic: USA (Arizona, California)
  • H. referens (Walker, 1852) - Oriental: India, Myanmar, Pakistan
  • H. robusta Zaitzev, 1966 - Palaearctic: Armenia, Azerbaijan, China, Gruzia, Kazakhstan, Kyrgyz Republic, Tajikistan, Turkmenistan, Uzbekistan
  • H. ruficollis (Bigot, 1892) - Neotropical: Argentina, Venezuela
  • H. scylla (Osten Sacken, 1887) - Nearctic: Mexico (Guanajuato, Sonora), USA (Arizona, Texas). Neotropical: Venezuela
  • H. semifucata (Hall, 1976) - Nearctic: USA (Arizona)
  • H. seminigra Loew, 1869 - Nearctic: Canada (Alberta, Saskatchewan), Mexico (Sonora), USA (Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, Utah, Washington). Neotropical: Puerto Rico (incl. H. eumenes)
  • H. sinuosa (Wiedemann, 1821) - Nearctic: USA
  • H. splendida Zaitzev, 1962 - Palaearctic: Kazakhstan, Kyrgyz Republic, Tajikistan, Turkmenistan
  • H. subarcuata Loew, 1871 - Palaearctic: Tajikistan, Tunisia
  • H. subvelutina Loew, 1871 - Palaearctic: Armenia, Azerbaijan, China (Shandong), Gruzia, Iran, Mongolia, Kazakhstan, Kyrgyz Republic, Tajikistan, Turkey, Turkmenistan, Uzbekistan
  • H. tarapacensis (Hall, 1976) - Neotropical: Chile
  • H. tenuirostris (Macquart, 1850) - Neotropical: Chile, Peru
  • H. translucens Ávalos-Hernández, 2009 - Nearctic: Mexico
  • H. tushetica Zaitzev, 1966 - Palaearctic: Armenia, Azerbaijan, China (Nei Monggol, Qinghai, Xinjiang), Gruzia, Iran
  • H. velutina (Meigen, 1820) - Oriental (Pakistan), Palaearctic
  • H. villeneuvi François, 1970 - Palaearctic: France, Greece, Italy
  • H. vockerothi François, 1969 - Palaearctic: Spain
  • H. webberi (Johnson, 1919) - Nearctic: Canada (Ontario, Quebec), USA (Connecticut, Kentucky, Massachusetts, Vermont, Wyoming)
  • H. wilcoxi (Painter, 1933) - Nearctic: USA (California, Washington)
  • H. yaqui (Painter in Painter & Painter, 1962) - Nearctic: Mexico (Sonora), USA (Arizona)

[4][5]

Extinct species

  • H. gabbroensis (Handlirsch, 1907) - (Miocene) Italy
  • H. provincialis (Handlirsch, 1907) - (Oligocene) France
  • H. tertiaria (Handlirsch, 1907) - (Oligocene) Germany

References

  1. 1 2 Hull, F.M. (1973). Bee flies of the world. The genera of the family Bombyliidae. Washington: Smithsonian Institution Press. pp. 687 pp. ISBN 0-87474-131-9.
  2. ↑ Evenhuis, N.L. (2020) A new genus for Painter & Painter’s Villa “celer”-group in the New World (Diptera: Bombyliidae). Zootaxa 4748(2): 296-314
  3. ↑ Ávalos-Hernández, O. (2009) A review of the North American species of Hemipenthes Loew, 1869 (Diptera: Bombyliidae) Zootaxa 2074. DOI: https://doi.org/10.11646/zootaxa.2074.1.1
  4. ↑ "World Catalog Family Bombyliidae Latreille (part 1 (Oligodraninae through Bombyliinae, p. 1-206)" (PDF). Bishop Museum. 1999. pp. 1–206. Retrieved 12 January 2009.
  5. ↑ "World Catalog Family Bombyliidae Latreille (part 1 (Oligodraninae through Bombyliinae, p. 1-206) - Revised Edition" (PDF). Bishop Museum. September 2003. pp. 1–206. Retrieved 12 January 2009.
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