Addition

Addition of species is defined by the disjoint union of sets, and corresponds to a choice between structures.^[6] For species F and G, define (F + G)[A] to be the disjoint union (also written "+") of F[A] and G[A]. It follows that (F + G)(x) = F(x) + G(x). As a demonstration, take E⁺ to be the species of non-empty sets, whose generating function is E⁺(x) = e^x − 1, and 1 the species of the empty set, whose generating function is 1(x) = 1. It follows that E = 1 + E⁺: in words, "a set is either empty or non-empty". Equations like this can be read as referring to a single structure, as well as to the entire collection of structures.

Multiplication

Multiplying species is slightly more complicated. It is possible to just take the Cartesian product of sets as the definition, but the combinatorial interpretation of this is not quite right. (See below for the use of this kind of product.) Rather than putting together two unrelated structures on the same set, the multiplication operator uses the idea of splitting the set into two components, constructing an F-structure on one and a G-structure on the other.^[7]

${\displaystyle (F\cdot G)[A]=\sum _{A=B+C}F[B]\times G[C].}$

This is a disjoint union over all possible binary partitions of A. It is straightforward to show that multiplication is associative and commutative (up to isomorphism), and distributive over addition. As for the generating series, (F · G)(x) = F(x)G(x).

The diagram below shows one possible (F · G)-structure on a set with five elements. The F-structure (red) picks up three elements of the base set, and the G-structure (light blue) takes the rest. Other structures will have F and G splitting the set in a different way. The set (F · G)[A], where A is the base set, is the disjoint union of all such structures.

The addition and multiplication of species are the most comprehensive expression of the sum and product rules of counting.

Composition

Composition, also called substitution, is more complicated again. The basic idea is to replace components of F with G-structures, forming (F∘G).^[8] As with multiplication, this is done by splitting the input set A; the disjoint subsets are given to G to make G-structures, and the set of subsets is given to F, to make the F-structure linking the G-structures. It is required for G to map the empty set to itself, in order for composition to work. The formal definition is:

${\displaystyle (F\circ G)[A]=\sum _{\pi \in P[A]}(F[\pi ]\times \prod _{B\in \pi }G[B]).}$

Here, P is the species of partitions, so P[A] is the set of all partitions of A. This definition says that an element of (F ∘ G)[A] is made up of an F-structure on some partition of A, and a G-structure on each component of the partition. The generating series is ${\displaystyle (F\circ G)(x)=F(G(x))}$.

One such structure is shown below. Three G-structures (light blue) divide up the five-element base set between them; then, an F-structure (red) is built to connect the G-structures.

These last two operations may be illustrated by the example of trees. First, define X to be the species "singleton" whose generating series is X(x) = x. Then the species Ar of rooted trees (from the French "arborescence") is defined recursively by Ar = X · E(Ar). This equation says that a tree consists of a single root and a set of (sub-)trees. The recursion does not need an explicit base case: it only generates trees in the context of being applied to some finite set. One way to think about this is that the Ar functor is being applied repeatedly to a "supply" of elements from the set — each time, one element is taken by X, and the others distributed by E among the Ar subtrees, until there are no more elements to give to E. This shows that algebraic descriptions of species are quite different from type specifications in programming languages like Haskell.

Likewise, the species P can be characterised as P = E(E⁺): "a partition is a pairwise disjoint set of nonempty sets (using up all the elements of the input set)". The exponential generating series for P is ${\displaystyle P(x)=e^{(e^{x}-1)}}$, which is the series for the Bell numbers.

Differentiation

Differentiation of species intuitively corresponds to building "structures with a hole", as shown in the illustration below.

Formally,^[9]

${\displaystyle (F')[A]=F[A\uplus \{\star \}],}$

where ${\displaystyle \star }$ is some distinguished new element not present in ${\displaystyle A}$.

To differentiate the associated exponential series, the sequence of coefficients needs to be shifted one place to the "left" (losing the first term). This suggests a definition for species: F' [A] = F[A + {*}], where {*} is a singleton set and "+" is disjoint union. The more advanced parts of the theory of species use differentiation extensively, to construct and solve differential equations on species and series. The idea of adding (or removing) a single part of a structure is a powerful one: it can be used to establish relationships between seemingly unconnected species.

For example, consider a structure of the species L of linear orders—lists of elements of the ground set. Removing an element of a list splits it into two parts (possibly empty); in symbols, this is L' = L·L. The exponential generating function of L is L(x) = 1/(1 − x), and indeed:

${\displaystyle {\frac {d}{dx}}{(1-x)}^{-1}={(1-x)}^{-2}.}$

The generalized differentiation formulas are to be found in a previous research by N. G. de Bruijn, published in 1964.

The species C of cyclic permutations takes a set A to the set of all cycles on A. Removing a single element from a cycle reduces it to a list: C' = L. We can integrate the generating function of L to produce that for C.

${\displaystyle C(x)=1+\int _{0}^{x}{\frac {dt}{1-t}}=1+\log {\frac {1}{1-x}}.}$

A nice example of integration of a species is the completion of a line (coordinatizated by a field) with the infinite point and obtaining a projective line.

Further operations

There are a variety of other manipulations which may be performed on species. These are necessary to express more complicated structures, such as directed graphs or bigraphs.

Pointing selects a single element in a structure.^[10] Given a species F, the corresponding pointed species F^• is defined by F^•[A] = A × F[A]. Thus each F^•-structure is an F-structure with one element distinguished. Pointing is related to differentiation by the relation F^• = X·F' , so F^•(x) = x F' (x). The species of pointed sets, E^•, is particularly important as a building block for many of the more complex constructions.

The Cartesian product of two species is a species which can build two structures on the same set at the same time. It is different from the ordinary multiplication operator in that all elements of the base set are shared between the two structures. An (F × G)-structure can be seen as a superposition of an F-structure and a G-structure. Bigraphs could be described as the superposition of a graph and a set of trees: each node of the bigraph is part of a graph, and at the same time part of some tree that describes how nodes are nested. The generating function (F × G)(x) is the Hadamard or coefficient-wise product of F(x) and G(x).

The species E^• × E^• can be seen as making two independent selections from the base set. The two points might coincide, unlike in X·X·E, where they are forced to be different.

As functors, species F and G may be combined by functorial composition: ${\displaystyle (F\,\Box \,G)[A]=F[G[A]]}$ (the box symbol is used, because the circle is already in use for substitution). This constructs an F-structure on the set of all G-structures on the set A. For example, if F is the functor taking a set to its power set, a structure of the composed species is some subset of the G-structures on A. If we now take G to be E^• × E^• from above, we obtain the species of directed graphs, with self-loops permitted. (A directed graph is a set of edges, and edges are pairs of nodes: so a graph is a subset of the set of pairs of elements of the node set A.) Other families of graphs, as well as many other structures, can be defined in this way.